The Natural World in Close Up

Collembola of Britain and Ireland
These maps record the presence of Collembola in the 10 km by 10 km Ordnance Survey Grid squares of Britain and Ireland (=hectads). For the maps of individual species, all published records are included as Green dots. Red dots indicate that presence has been confirmed by examination of microscope slides in the Natural History Museum, London (NHML), specimens in my reference collection, or material sent to me by recorders. Note that Orkney and Shetland have been relocated from their true geographic positions. The maps were produced with the DMAP computer program developed by Alan Morton [http://www.dmap.co.uk].
     Dots will be added to the existing maps as new localities are discovered. The date on which records were most recently added is shown in the title adjacent to the species name on each map. Each species has been allocated a code number that will never change, and a five letter abbreviation usually made up of the first two letters of the genus in capitals and first three letters of the species name in lower case. For example, the code and abbreviation for Brachystomella parvula is 001/BRpar). The five letter abbreviation is subject to alteration when species are moved between genera but the fixed number code will ensure that species can be tracked over time, irrespective of name changes.
     The checklist below includes changes arising from recent taxonomic decisions, new synonyms, and the discovery of several misidentifications in the NHML Collembola collections. Several 'species' are undoubtedly complexes of two or more species. However, I have decided to adopt a conservative approach until fresh UK/Eire material can be examined to resolve some of the more contentious species boundaries. Synonyms (*) are listed below each species name.
     In most cases, genus and species names have been updated to conform with those used in the publications listed below. The arrangement of Families and Subfamilies follows the scheme of Deharveng (2004).

Babenko et al. (1994). Collembola of Russia and Adjacent Countries: Family Hypogastruridae, Nauka, Moscow.
Bretfeld, G. (1999). Synopses on Palaearctic Collembola: Symphypleona. Abhandlungen und Berichte des Naturkundemuseums Görlitz, 71, (1), 1-318.
Deharveng, L. (2004). Recent advances in Collembola systematics. Pedobiologia, 48, 415-433.
Fjellberg (1998). The Collembola of Fennoscandia and Denmark Part 1: Poduromorpha, Fauna Entomologica Scandinavica, Vol. 35, Brill, Leiden.
Jordana, R., Arbea, J.I., Simon, C. & Lucianez, M.J. (1997). Collembola, Poduromorpha. In Fauna Ibérica, Volume 8 (ed. M.A. Ramos). Museo Nacional de Ciencas Naturales, CSIC, Madrid, 807pp.
Pomorski, J. (1998). Onychiurinae of Poland (Collembola: Onychiuridae). Polish Taxonomical Society, Wrocklaw, 201pp.
Potapov, M. (2001). Synopses on Palaearctic Collembola: Isotomidae. Abhandlungen und Berichte des Naturkundemuseums Görlitz, 73, (2), 1-603.
Thibaud, J.M., Schulz, H.J. & da Gama Assalino, M.M. (2004). Synopses on Palaearctic Collembola: Hypogastruridae. Abhandlungen und Berichte des Naturkundemuseums Görlitz, 75, (2), 1-287.
Zimdars, B. & Dunger, W. (1994). Synopses on Palaearctic Collembola: Tullbergiinae. Abhandlungen und Berichte des Naturkundemuseums Görlitz, 68, (4), 1-71.

*** Coverage and Species Richness Maps ***

Click here for a map of the hectads from which at least one species of Collembola has been recorded (Coverage Map)
Click here for a map showing the number of species recorded from each hectad (Species Richness Map)

*** Status ***

VERY COMMON = present in more than 50 hectads
COMMON = present in 26 to 50 hectads
MODERATELY COMMON = present in 11 to 25 hectads
SCARCE = present in 6 to 10 hectads
RARE = present in 5 or fewer hectads
[P] = Photograph(s) included of whole animal and/or diagnostic features

There are 89 VERY COMMON or COMMON species; these are shown in BOLD UPPER CASE text.
There are 160 MODERATELY COMMON, SCARCE or RARE species; these are shown in bold lower case text.
There are 140 species shown in unemphasised lower case text for which there are no definite records supported by specimens and which are unlikely to be present in UK/Eire. Maps are included of literature records for some of these species.

*** Click on the species name below to access the map ***


Class Collembola

Order Poduromorpha

Superfamily Hypogastruroidea

Family Hypogastruridae

Genus Ceratophysella Börner 1932
(002/CEarm) Ceratophysella armata (Nicolet, 1842) [NO CONFIRMED UK/EIRE RECORDS]
(003/CEben) CERATOPHYSELLA BENGTSSONI (Ågren, 1904) [COMMON, WIDESPREAD] [P]
          *angularis (Bagnall, 1949)
          *collingei (Bagnall, 1949)
(004/CEden) CERATOPHYSELLA DENTICULATA (Bagnall, 1941) [VERY COMMON, WIDESPREAD] [P]
         *distinguenda (Bagnall, 1941)
(005/CEeng) Ceratophysella engadinensis (Gisin, 1949) [RARE] [P]
(006/CEgib) Ceratophysella gibbosa (Bagnall, 1940) [MODERATELY COMMON] [P]
(007/CEgra) Ceratophysella granulata Stach, 1949 [NO CONFIRMED UK/EIRE RECORDS]
(008/CElon) Ceratophysella longispina (Tullberg, 1876) [not (Willem, 1902)] [ARCTIC SPECIES (UK/EIRE SPECIMENS ARE
          OTHER SPECIES OF Ceratophysella)]
(009/CEruf) Ceratophysella rufescens (Nicolet, 1842) [SINGLE UK RECORD WAS PROBABLY Ceratophysella bengtssoni]
(010/CEsco) CERATOPHYSELLA SCOTICA (Carpenter & Evans, 1899) [COMMON, WIDESPREAD IN BOGS AND WET
          HABITATS] [P]
          *gotoi (Murphy, 1960)
(011/CEsig) Ceratophysella sigillata (Uzel, 1891) [UK SPECIMENS ARE Ceratophysella bengtssoni]
(012/CEsuc) Ceratophysella succinea (Gisin, 1949) [NO CONFIRMED UK/EIRE RECORDS]

Genus Choreutinula Paclt 1944
(012.5/CHine) Choreutinula inermis (Tullberg, 1871) [NO CONFIRMED UK/EIRE RECORDS]

Genus Hypogastrura Bourlet 1839
(013/HYass) Hypogastrura assimilis Krausbauer, 1898 [RARE] [P]
(014/HYbur) Hypogastrura burkilli (Bagnall, 1940) [RARE] [P]
(014.5/HYdis) Hypogastrura distincta (Axelson, 1902) [RARE] [P]
(015/HYele) Hypogastrura elegans (Parfitt, 1891) [SINGLE UK RECORD IS A Ceratophysella (SPECIES UNKNOWN)]
(016/HYlap) Hypogastrura lapponica (Axelson, 1902) [BOREAL SPECIES, UNLIKELY TO BE PRESENT]
          *frigida (Axelson, 1905)
          *torneensis (Wahlgren, 1906)
(017/HYlit) Hypogastrura litoralis (Axelson, 1907) [SCARCE] [P]
          *browni (Bagnall, 1940)
(018/HYman) HYPOGASTRURA MANUBRIALIS (Tullberg, 1869) [COMMON, WIDESPREAD] [P]
          *mucronalis (Bagnall, 1941)
          *neglecta (Börner, 1901) [VARIETY OF Hypogastrura manubrialis WITH NO ANAL SPINES]
          *schoetti (Reuter, 1895)
(019/HYpac) Hypogastrura packardi (Folsom, 1902) [NO CONFIRMED UK/EIRE RECORDS]
(020/HYpur) HYPOGASTRURA PURPURESCENS (Lubbock, 1867) [VERY COMMON, WIDESPREAD] [P]
          *propinqua (Carpenter & Evans, 1904)
          *pseudo-purpurascens Womersley, 1928
(022/HYsah) Hypogastrura sahlbergi (Reuter, 1895) [SCARCE] [P]
          *schaefferi (Carl, 1901)
          *schneideri (Schäffer, 1896)
          *strenua (Brown, 1923)
(023/HYser) Hypogastrura serrata (Ågren, 1904) [NO CONFIRMED UK/EIRE RECORDS]
          *ripperi Gisin, 1952
(024/HYsoc) Hypogastrura socialis (Uzel, 1891) [NO CONFIRMED UK/EIRE RECORDS]
          *mitis (Ågren, 1904)
(025/HYtul) Hypogastrura tullbergi (Schäffer, 1900) [ARCTIC SPECIES (UK/EIRE SPECIMENS ARE OTHER SPECIES
          OF Hypogastrura)]
          *dubia (Tullberg, 1876) [dubia of Templeton, 1835 was a juvenile Podura aquatica]
(026/HYver) Hypogastrura vernalis (Carl, 1901) [RARE] [P]
          *nivalis (Ågren, 1903)
          *reuteri (Ågren, 1903)
(027/HYvia) HYPOGASTRURA VIATICA (Tullberg, 1872) [VERY COMMON, WIDESPREAD] [P]
          *pseudoviatica (Bagnall, 1941)
          *subviatica (Bagnall, 1941)

Genus Mesogastrura Bonet 1930
(028/MElib) Mesogastrura libyca (Caroli, 1914) [RARE] [P]

Genus Schaefferia Absolon 1900
(030/SCemu) SCHAEFFERIA EMUCRONATA Absolon, 1900 'group' [VERY COMMON, WIDESPREAD, OFTEN IN
          CAVES] [P]
(030.5/SClin) Schaefferia lindbergi da Gama, 1962 [MAPPED UNDER Schaefferia emucronata 'group']
(032/SCpou) Schaefferia pouadensis Delamare Deboutteville, 1945 [MAPPED UNDER Schaefferia emucronata 'group']
(032.2/SCsex) Schaefferia sexoculata (Gisin, 1947) [MAPPED UNDER Schaefferia emucronata 'group']
(032.5/SCwil) Schaefferia willemi (Bonet, 1930) [MAPPED UNDER Schaefferia emucronata 'group']
          *cavernicola (Womersley, 1930)
          *longispina (Willem, 1902) [not (Tullberg, 1876)]
          *octoculata (Womersley, 1930)

Genus Schoettella Schäffer 1896
(033/SHunu) Schoettella ununguiculata (Tullberg, 1869) [RARE] [P]
          *tullbergi (Lubbock, 1873)

Genus Willemia Börner 1901
(034/WLano) WILLEMIA ANOPHTHALMA Börner, 1901 [COMMON, WIDESPREAD] [P]
(036/WLbud) Willemia buddenbrocki Hüther, 1959 [RARE] [P]
(037/WLden) Willemia denisi Mills, 1932 [SCARCE] [P]
          *aspinata Stach, 1949
(038/WLint) Willemia intermedia Mills, 1934 [RARE] [P]
(038.5/WLmul) Willemia multilobata Gers & Deharveng, 1985 [RARE]
(039/WLsca) Willemia scandinavica  Stach, 1949 [RARE] [P]

Genus Xenylla Tullberg 1869
(040/XLaca) Xenylla acauda Gisin, 1947 [SCARCE, SW BIAS TO DISTRIBUTION] [P]
(041/XLboe) Xenylla boerneri Axelson, 1905 [MODERATELY COMMON, WIDESPREAD] [P]
(042/XLbre) Xenylla brevicauda Tullberg, 1869 [NO CONFIRMED UK/EIRE RECORDS]
(043/XLcor) Xenylla corticalis Börner, 1901 [RARE] [P]
(044/XLgri) XENYLLA GRISEA Axelson, 1900 [COMMON, WIDESPREAD] [P]
(045/XLhum) XENYLLA HUMICOLA (O. Fabricius, 1780) [COMMON, WIDESPREAD, MAINLY LITTORAL] [P]
(046/XLlon) Xenylla longispina Uzel, 1891 [DOUBTFUL SPECIES]
(047/XLmar) Xenylla maritima Tullberg, 1869 [MODERATELY COMMON, MAINLY LITTORAL] [P]
(048/XLmuc) Xenylla mucronata Axelson, 1903 [NO CONFIRMED UK/EIRE RECORDS]
(049/XLtul) Xenylla tullbergi Börner 1903 [RARE] [P]
(050/XLwel) Xenylla welchi Folsom, 1916 [SCARCE, INTRODUCED WITH MUSHROOMS?] [P]
          *subwelchi Denis, 1924
(051/XLxav) Xenylla xavieri Gama, 1959 [SCARCE] [P]

Superfamily Neanuroidea

Family Brachystomellidae

Genus Brachystomella Ågren 1903
(001/BRpar) BRACHYSTOMELLA PARVULA (Schäffer, 1896) [COMMON, WIDESPREAD] [P]
          *crassicornis (Schött, 1902)
          *maritima Ågren, 1903
          *media (Axelson, 1900)
          *wahlgreni (Denis, 1924)

Family Neanuridae

Subfamily Frieseinae

Genus Friesea Dalle Torre 1895
(052/FRacu) Friesea acuminata (Denis, 1925) [SCARCE, LITTORAL] [P]
          *thalassophila (Bagnall, 1939)
(053/FRafu) Friesea afurcata (Denis, 1926) [SINGLE EIRE RECORD NEEDS CONFIRMATION]
(054/FRcla) FRIESEA CLAVISETA Axelson, 1900 [COMMON, WIDESPREAD] [P]
(055/FRemu) Friesea emucronata (Stach, 1922) [FEW RECORDS, PROBABLY ALL Friesea truncata]
(056/FRmir) FRIESEA MIRABILIS (Tullberg, 1871) [VERY COMMON, WIDESPREAD] [P]
          *poseidonis Bagnall, 1939
(057/FRtru) Friesea truncata Cassagnau, 1958 [MODERATELY COMMON, WIDESPREAD] [P]

Subfamily Neanurinae

Genus Lathriopyga Caroli 1910
(058/LAlon) Lathriopyga longiseta (Caroli, 1912) [SCARCE, SOUTHWEST ENGLAND ONLY] [P]
          *hystrix (Bagnall, 1940)

Genus Monobella Cassagnau 1979
(059/MOgra) Monobella grassei (Denis, 1923) [MODERATELY COMMON IN SOUTHERN ENGLAND] [P]

Genus Neanura MacGillivray 1893
(060/NNmus) NEANURA MUSCORUM (Templeton, 1835) [VERY COMMON, WIDESPREAD] [P]
          *tuberculata (Nicolet. 1842)

Genus Paranura Axelson 1902
(061/PUsex) Paranura sexpunctata Axelson, 1902 [NO CONFIRMED UK/EIRE RECORDS]

Genus Yuukianura Yosii 1955
(062/YUaph) Yuukianura aphoruroides (Yosii, 1953) [RARE, INTRODUCED, PROBABLY EXTINCT IN UK/EIRE]

Subfamily Pseudachorutinae

Genus Anurida Laboulbene 1865
(064/ANden) Anurida denisi (Bagnall, 1939) [SCARCE, LITTORAL] [P]
          *hibernica (Bagnall, 1941)
          *trioculata Denis, 1922 [not (Kinoshita, 1916)]
(065/ANell) Anurida ellipsoides Stach, 1949 [NO CONFIRMED UK/EIRE RECORDS]
(066/ANgra) ANURIDA GRANARIA (Nicolet, 1847) [VERY COMMON. WIDESPREAD] [P]
          *bagnalli Stach, 1949
          *comellinii Gisin, 1949
(068/ANmar) ANURIDA MARITIMA (Guérin, 1839) [VERY COMMON, LITTORAL] [P]
          *bisetosa Bagnall, 1949 [possibly a good species]
          *megalops Bagnall, 1949
(070/ANtha) Anurida thalassophila (Bagnall, 1939) [MODERATELY COMMON, LITTORAL] [P]
          *remyi Denis, 1947
(071/ANtul) Anurida tullbergi Schött, 1891 [MODERATELY COMMON, WIDESPREAD] [P]
          *crassicornis (Reuter & Reuter, 1880)

Genus Anuridella Willem 1906
(072/ADcal) Anuridella calcarata Denis, 1925 [NO CONFIRMED UK/EIRE RECORDS]
(073/ADimm) Anuridella immsiana Bagnall, 1939 [RARE, LITTORAL] [P]
(074/ADmar) Anuridella marina Willem, 1906 [SCARCE, LITTORAL] [P]
          *hintoni Bagnall, 1949
(075/ADsub) Anuridella submarina Bagnall, 1934 [RARE, LITTORAL] [P]
          *germanica Gisin, 1951

Genus Micranurida Börner 1901
(076/MIfor) Micranurida forsslundi Gisin, 1949 [RARE] [P]
          *conjuncta Murphy, 1960
(077/MIgra) Micranurida granulata (Agrell, 1943) [RARE]
(078/MIpyg) MICRANURIDA PYGMAEA Börner, 1901 [COMMON, WIDESPREAD] [P]
(079/MIsen) Micranurida sensillata (Gisin, 1953) [RARE] [P]

Genus Pseudachorutella Stach 1949
(080/PLasi) Pseudachorutella asigillata (Börner, 1901) [MODERATELY COMMON, WIDESPREAD] [P]
          *silvaticus (Agren, 1903)
(081/PLcla) Pseudachorutella clavata (Börner, 1901) [DOUBTFUL SPECIES]

Genus Pseudachorutes Tullberg 1871
(082/PCboe) Pseudachorutes boerneri Schött, 1902 [NO CONFIRMED UK/EIRE RECORDS]
          *forsslundi Agrell, 1943
(083/PCcor) Pseudachorutes corticicolus (Schäffer, 1896) [RARE]
(084/PCdub) Pseudachorutes dubius Krausbauer, 1898 [NO CONFIRMED UK/EIRE RECORDS]
          *lapponicus Ågren, 1904
(085/PCpar) Pseudachorutes parvulus Börner, 1901 [NO CONFIRMED UK/EIRE RECORDS]
(086/PCsub) Pseudachorutes subcrassus Tullberg, 1871 'group' [MODERATELY COMMON (SPECIES COMPLEX?)] [P]
          *tuberculatus Schött, 1902

Family Odontellidae

Genus Odontella Schäffer 1897
(087/ODemp) Odontella empodialis Stach, 1934 [SINGLE WELSH RECORD NEEDS CONFIRMATION]
(088/ODlam) Odontella lamellifera (Axelson, 1930) 'group' [RARE (SPECIES COMPLEX?)] [P]
          *suecica Ågren, 1903

Genus Xenyllodes Axelson 1903
(089/XOarm) Xenyllodes armatus Axelson, 1903 [SCARCE] [P]

Superfamily Onychiuroidea

Family Onychiuridae

[NOTE: Although splitting the Onychiuridae into several genera can be criticised, it does have the advantage of separating out the definite species groups within the old definitions of Protaphorura (post-antennal organ (PAO) with simple vesicles) and Onychiurus (PAO with complex vesicles). Generic boundaries are by no means settled and further changes are likely in the future. The genera used in this list are those of Pomorski (1988) in his Synopsis on the Onychiurinae of Poland. The list below represents my first attempt to clarify a messy situation by not confirming the presence of a species on the checklist ('NO CONFIRMED UK/EIRE RECORDS'] unless it can be supported by a specimen in good condition. It is my opinion that several of these are not good species (especially Protaphorura sp. described by Gisin). The checklist is by no means 'cast in stone' and undoubtedly contains errors. I am certain that some of the species described by Gisin had already been seen (and in some cases described) by Bagnall. However, the descriptions of both authors lack the detail expected of modern accounts and most of their type specimens are in very poor condition. A thorough revision is needed of all Bagnall's and Gisin's 'species'. I am grateful to Arne Fjellberg and Jacek Pomorski for their valuable comments on this list, although they do not agree entirely with my conclusions for which I am fully responsible.]

Genus Allonychiurus Yoshii 1995
(101/AOedi) Allonychiurus edinensis (Bagnall, 1935) [MODERATELY COMMON] [P]
          *spinularius (Gisin, 1952)
(113/AOspi) Allonychiurus spinosus (Bagnall, 1949) [NO CONFIRMED UK/EIRE RECORDS]

Genus Bionychiurus Pomorski 1996
(109.5/BInor) Bionychiurus normalis (Gisin, 1949) [NO CONFIRMED UK/EIRE RECORDS, MAY TURN UP IN
          GLASSHOUSES]

Genus Deharvengiurus Weiner 1996
(097/DHarg) Deharvengiurus argus (Denis, 1925) [RARE, TRESCO ONLY]

Genus Detriturus Pomorski 1988
(106/DTjub) Detriturus jubilarius (Gisin, 1957) [RARE] [P]

Genus Deuteraphorura Absolon 1901
(098/DUine) DEUTERAPHORURA INERMIS (Tullberg, 1871) [COMMON, WIDESPREAD] [P]
          *cebennaria (Gisin, 1956)
          *dunaria (Gisin, 1956)
          *fimetaria (Linnaeus, 1767) [not 1758] [INVALID NAME]
          *imperfectus (Denis, 1938)
          *pygmaea (Bagnall, 1937)
          *wrighti (Carpenter, 1895)
(109/DUner) Deuteraphorura nervosa (Stach, 1954) [NO CONFIRMED UK/EIRE RECORDS]
(111/DUsco) Deuteraphorura scotaria (Gisin, 1954) [SCARCE] [P]
(116/DUvar) Deuteraphorura variabilis (Stach, 1954) [NO CONFIRMED UK/EIRE RECORDS]

Genus Kalaphorura Absolon 1901
(092/KUbea) Kalaphorura bearei (Bagnall, 1937) [ALL SPECIMENS ARE JUVENILES OF A SPECIES OF Onychiurus]
(093/KAbur) KALAPHORURA BURMEISTERI (Lubbock, 1873) [COMMON, MAINLY IN SOUTH] [P]
          *tuberculata (Moniez, 1891)

Genus Megaphorura Fjellberg 1998
(118/MAarc) Megaphorura arctica (Tullberg, 1876) [RARE, SKOMER ISLAND ONLY] [P]

Genus Micraphorura Bagnall 1949
(090/MCabs) Micraphorura absoloni (Börner, 1901) [MODERATELY COMMON (NOT IRELAND)] [P]
          *affinis (Ågren, 1903)
          *handschini (Bagnall, 1949)

Genus Micronychiurus Bagnall 1949
(108/MYmin) Micronychiurus minutus (Denis, 1932) [NO CONFIRMED UK/EIRE RECORDS]

Genus Oligaphorura Bagnall 1949
(091/OLsch) OLIGAPHORURA SCHOETTI (Lie Pettersen, 1897) [COMMON IN CAVES] [P]

Genus Onychiurus Gervais 1841
(095/ONamb) ONYCHIURUS AMBULANS (Linnaeus, 1758) [VERY COMMON, WIDESPREAD] [P]
          *circulans Gisin, 1952
          *laminatipes Bagnall, 1937
(096/ONara) Onychiurus arans Gisin, 1952 [NO CONFIRMED UK/EIRE RECORDS]
[Onychiurus circulans see Onychiurus ambulans]
(099/ONdis) Onychiurus dissimulans Gisin, 1952 [NO CONFIRMED UK/EIRE RECORDS]
(102/ONfid) Onychiurus fimetarioides Denis, 1938 [NO CONFIRMED UK/EIRE RECORDS]
          *pseudofimetarius Handschin 1920 [not Folsom, 1917]
[Onychiurus fimetarius see Deuteraphorura inermis]
(110/ONrec) Onychiurus rectospinatus Stach, 1922 [NO CONFIRMED UK/EIRE RECORDS]
(115/ONsub) Onychiurus subambulans Denis, 1935 [NO CONFIRMED UK/EIRE RECORDS]

Genus Orthonychiurus Stach 1954
(104/OTfol) Orthonychiurus folsomi (Schäffer, 1900) [RARE, GLASSHOUSES ONLY] [P]
          *gotoi (Chowdhury, 1958)
          *musae (Selga, 1962)
          *sinensis (Stach, 1954)
(114/OTsta) Orthonychiurus stachianus (Bagnall, 1939) [NO CONFIRMED UK/EIRE RECORDS]
          *pseudostachianus (Gisin, 1956)
          *stachi (Denis, 1938) [not Bagnall, 1935]

Genus Protaphorura Absolon 1901

[NOTE: Numerous species within the Protaphorura armata 'group' have been described (simple vesicles in PAO, furca present as vestigial flap). All the 'species' recorded in the literature are listed below. Synonyms are based on the checklist of Kloet & Hincks (1964), Fjellberg (1998) and Pomorski (1998), and some of my own observations. However, after examining more than 1000 specimens of Protaphorura in the NHML slide collection (including all of Bagnall's surviving material), I have come to the conclusion that it is impossible to reliably allocate more than three names to these UK/Eire specimens on the basis of characters used by Gisin and others. The pseudocelli formula, presence or absence of seta s' on abd5, and the degree of convergence of lines drawn through the two pairs of setae anterior to the anal spines are too variable. That is not to say that there are not more than three good species of Protaphorura in UK/Eire but that the morphological characters that are currently used to separate them are not 100% reliable. Bring on the molecules!]

(117/PRalb) Protaphorura alborufescens (Vogler, 1895) [UK/EIRE SPECIMENS ARE Protaphorura aurantiaca]
(119/PRarm) PROTAPHORURA ARMATA (Tullberg, 1869) [VERY COMMON, WIDESPREAD] [P]
          *humata (Gisin, 1952)
          *nemorata (Gisin, 1952)
          *subaequalis (Bagnall, 1937)
          *waterstoni (Bagnall, 1937)
(120/PRaur) PROTAPHORURA AURANTIACA (Ridley, 1880) [VERY COMMON, WIDESPREAD] [P]
          *daviesi (Bagnall, 1935)
          *flavescens (Bagnall, 1935)
          *flavidula (Bagnall, 1939)
          *lata (Gisin, 1956)
          *magnicornis (Bagnall, 1937)
          *prolata (Gisin, 1956)
          *sublata (Gisin, 1957)
          *uliginata (Gisin, 1952)
(121/PRbag) Protaphorura bagnalli (Salmon, 1959) [HOLOTYPE IS JUVENILE, PROBABLE JUVENILE SYNONYM OF
     Protaphorura armata]
(122/PRbic) Protaphorura bicampata (Gisin, 1956) [NO CONFIRMED UK/EIRE RECORDS]
(123/PRcal) Protaphorura caledonica (Bagnall, 1935) [PROBABLE JUNIOR SYNONYM OF Protaphorura armata]
(124/PRcam) Protaphorura campata (Gisin, 1952) [NO CONFIRMED UK/EIRE RECORDS]
          *procampata (Gisin, 1956)
(128/PRfim) Protaphorura fimata (Gisin, 1952) [UK/EIRE SPECIMENS ARE Protaphorura aurantiaca]
(132/PRmac) Protaphorura macfadyeni (Gisin, 1953) [RARE, LITTORAL] [P]
(137/PRmer) Protaphorura meridiata (Gisin, 1952) [NO CONFIRMED UK/EIRE RECORDS]
          *pulvinata (Gisin, 1954)
(139/PRoct) Protaphorura octopunctata (Tullberg, 1876) [UK/EIRE SPECIMENS ARE Protaphorura armata OR Protaphorura
          aurantiaca]
(140/PRpan) Protaphorura pannonica (Haybach 1960) [NO CONFIRMED UK/EIRE RECORDS]
          trinotata (Gisin 1961)
          pseudocellata (Naglitsch, 1962)
(151/PRpse) Protaphorura pseudovanderdrifti (Gisin, 1957) [NO CONFIRMED UK/EIRE RECORDS]
          *s-vontoernei (Gisin, 1957)
(145/PRqua) Protaphorura quadriocellata (Gisin 1947) [UK/EIRE SPECIMENS ARE Protaphorura aurantiaca]
(146/PRsta) Protaphorura stachi (Bagnall, 1935) [not (Denis, 1938)] [HOLOTYPE IS JUVENILE; PROBABLE JUNIOR
          SYNONYM OF Protaphorura armata]
(150/PRsub) Protaphorura subuliginata (Gisin, 1956) [UK/EIRE SPECIMENS ARE Protaphorura armata OR Protaphorura
          aurantiaca]
          *subarmata (Gisin, 1957)
(153/PRtri) Protaphorura tricampata (Gisin, 1956) [NO CONFIRMED UK/EIRE RECORDS]
(155/PRtul) Protaphorura tullbergi [PROBABLE JUNIOR SYNONYM OF Protaphorura armata]

Genus Supraphorura Stach 1954
(130/SRfur) SUPRAPHORURA FURCIFERA (Börner, 1901) [COMMON, WIDESPREAD] [P]
          *celtica (Bagnall, 1937)
          *furcifer (Axelson, 1905)
          *?scotica (Bagnall, 1935)

Genus Tantulonychiurus Pomorski 1996
(116.5/TAvol) Tantulonychiurus volinensis (Szeptycki, 1964) [NO UK/EIRE RECORDS, MAY TURN UP IN SAND DUNES]

Genus Thalassaphorura Bagnall 1949
(125/THdeb) THALASSAPHORURA DEBILIS (Moniez, 1890) [COMMON, LITTORAL] [P]
          *imminuta (Bagnall, 1937)
          *thalassophila (Bagnall, 1937)
(126/THenc) Thalassaphorura encarpata (Denis, 1931) [RARE, GLASSHOUSES ONLY] [P]
          *hortensis (Gisin, 1949)
          *parthenogenetica (Choudhuri, 1958)
(127/THeva) Thalassaphorura evansi [PROBABLE JUNIOR SYNONYM OF Protaphorura debilis]
          *moniezi (Bagnall, 1935)
(131/THhal) Thalassaphorura halophila (Bagnall, 1937) [SCARCE, LITTORAL] [P]
          *littoralis (Bagnall, 1937)

Family Tullbergiidae

Genus Mesaphorura Börner 1901
(158/MSatl) Mesaphorura atlantica (Rusek, 1979) [STATUS UNCLEAR, RARE?]
(158.2/MScri) Mesaphorura critica Ellis, 1976 [RARE]
(158.5/MShyl) Mesaphorura hylophila (Rusek, 1982) [RARE] [P]
(159/MSiow) Mesaphorura iowensis (Mills, 1932) [NO CONFIRMED UK/EIRE RECORDS]
(160/MSkra) Mesaphorura krausbaueri Börner, 1901 [MODERATELY COMMON?, SEE MAP FOR COMMENTS] [P]
          *thalassophila Bagnall, 1937
(161/MSmac) MESAPHORURA MACROCHAETA Rusek, 1976 [COMMON, WIDESPREAD] [P]
(161.2/MSpet) Mesaphorura petterdassi (Fjellberg, 1988) [RARE]
(161.5/MSpon) Mesaphorura pongei Rusek, 1982 [RARE]
(162/MSsyl) Mesaphorura sylvatica (Rusek, 1971) [RARE]
(163/MSten) Mesaphorura tenuisensillata Rusek, 1974 [RARE, HIGH MOUNTAINS ONLY] [P]
(164/MSyos) Mesaphorura yosiii (Rusek, 1967) [RARE, ACIDIC HABITATS] [P]

Genus Metaphorura Stach 1954
(165/MTaff) METAPHORURA AFFINIS (Börner, 1902) [COMMON, WIDESPREAD] [P]
          *bipartita (Handschin, 1920)
          *boerneri Bagnall, 1936

Genus Neonaphorura Bagnall 1935
166/NPdub) Neonaphorura duboscqi (Denis, 1932) [RARE] [P]
          *anglicana Bagnall 1936

Genus Neotullbergia Bagnall 1935
(167/NTcra) Neotullbergia crassicuspis (Gisin, 1944) [RARE] [P]
(168/NTram) Neotullbergia ramicuspis (Gisin 1953) [RARE] [P]
(169/NTtri) Neotullbergia tricuspis (Börner, 1902) [RARE] [P]
          *laingi Bagnall, 1936

Genus Paratullbergia Womersley 1930
(170/PTcal) PARATULLBERGIA CALLIPYGOS (Börner, 1902) [VERY COMMON, WIDESPREAD] [P]
          *carpenteri Bagnall, 1935
          *concolor Womersley 1930
          *womersleyi Bagnall, 1935
(173/PTmac) Paratullbergia macdougalli Bagnall, 1936 [SCARCE] [P]

Genus Stenaphorura Absolon 1900
(176/SUden) STENAPHORURA DENISI (Bagnall, 1935) [COMMON, WIDESPREAD] [P]
          *parisi (Denis, 1943)
(175/SUjap) Stenaphorura japygiformis Absolon, 1900 [UNLIKELY TO BE PRESENT]
(177/SUlub) Stenaphorura lubbocki (Bagnall, 1935) [MODERATELY COMMON, WIDESPREAD] [P]
(178/SUqua) STENAPHORURA QUADRISPINA (Börner, 1901) [COMMON, WIDESPREAD] [P]
          *absoloni (Bagnall, 1936)
          *axelsoni (Bagnall, 1935)

Incertae sedis (i.e. taxonomic position not clear)

Family Poduridae

Genus Podura Linnaeus 1758
(179/PDaqu) PODURA AQUATICA Linnaeus, 1758 [COMMON, WIDESPREAD ON SURFACES OF PONDS
          AND LAKES] [P]


Order Entomobryomorpha

Superfamily Entomobryoidea

Family Cyphoderidae

Genus Cyphoderus Nicolet 1842
(180/CYalb) CYPHODERUS ALBINUS Nicolet, 1842